1. School of Biological Sciences Seoul National University Gwanak‐ro Seoul 08826 Korea
2. National Institute of Horticultural & Herbal Science Rural Development Administration Wanju 55365 Korea
3. Department of Medical & Biological Sciences The Catholic University of Korea Jibong‐ro Bucheon‐si Gyeonggi‐do 14662 Korea
4. Department of Biological Sciences National University of Singapore 14 Science Drive 4 Singapore 117543 Singapore
5. Department of Agriculture, Forestry and Bioresources Seoul National University Gwanak‐ro Seoul 08826 Korea
6. Plant Genomics and Breeding Institute Seoul National University Gwanak‐ro Seoul 08826 Korea