1. The Swire Institute of Marine Science and Division of Ecology & Biodiversity, The School of Biological Sciences, The University of Hong Kong, Pokfulam Road, Hong Kong, SAR, China
2. Academy Sea Environment, Lungomare dei navigatori 44, 58019 Monte Argentario, Italy
3. School of Biological Sciences, Royal Holloway, University of London, Egham, TW20 0EX, UK