1. Laboratoire Ecologie des Systèmes Marins Côtiers UMR 5119 CNRS, IRD, IFREMER, UM2, UM1 cc 093, Place E. Bataillon 34095 Montpellier Cedex 5 France
2. Laboratoire Ecosystèmes Marins Exploités UMR 212 IRD, IFREMER, UM2 avenue Jean Monnet BP171 34203 Sète Cedex France
3. Rui Nabeiro Biodiversity Chair CIBIO, University of Évora Casa Cordovil, 2º Andar, Rua Dr. Joaquim Henrique da Fonseca 7000‐890 c Portugal
4. Departamento de Biodiversidad y Biologia Evolutiva Museo Nacional de Ciencias Naturales, CSIC C/José Gutiérrez Abascal, 2 28006 Madrid Spain
5. Center for Macroecology, Evolution and Climate University of Copenhagen Universitetsparken 15 2100 Copenhagen Denmark
6. ARC Centre of Excellence for Coral Reef Studies James Cook University Townsville Qld 4811 Australia