1. Key Laboratory of Plant Resources Conservation and Sustainable Utilization, South China Botanical Garden, Chinese Academy of Sciences Guangzhou 510650 China
2. University of Chinese Academy of Sciences Beijing 100049 China
3. Shenzhen Academy of Metrology and Quality Inspection Shenzhen 518109 China
4. Germplasm Bank of Wild Species, Kunming Institute of Botany, Chinese Academy of Sciences Kunming 650201 China
5. Department of Geosciences and Natural Resource Management University of Copenhagen 1958 Frederiksberg Denmark
6. US National Herbarium, Department of Botany Smithsonian Institution‐NMNH Washington DC 20560 USA
7. Herbarium, Royal Botanic Gardens, Kew, Richmond Surrey TW9 3AE UK
8. School of Pharmaceutical Science, Guangzhou University of Chinese Medicine Guangzhou 510006 China