1. Department of Natural History Sciences Graduate School of Science Hokkaido University Sapporo 060‐0810 Japan
2. Asian Natural Environmental Science Center The University of Tokyo 1‐1‐1 Yayoi Bunkyo Tokyo113‐8657 Japan
3. Research Institute for Marine Fisheries 222 Le Hai Hai Phong Vietnam
4. Department of Biological Sciences Faculty of Science Hokkaido University Sapporo 060‐0810 Japan