1. Freshwater Biological Laboratory Department of Biology University of Copenhagen Universitetsparken 4, 3rd floor 2100 Copenhagen Denmark
2. UWA School of Agriculture and Environment Faculty of Science The University of Western Australia Crawley WA 6009 Australia
3. Graduate School of Bioagricultural Sciences Nagoya University Furo‐cho Chikusa Nagoya 464‐8601 Japan
4. Forage Crop Research Division Institute of Livestock and Grassland Science NARO 768 Senbonmatsu Nasushiobara Tochigi 329‐2793 Japan