1. Coastal Ecosystems Research Station of the Yangtze River Estuary, Ministry of Education Key Laboratory for Biodiversity Science and Ecological Engineering, The Institute of Biodiversity Science, Fudan University, 220 Handan Road, Shanghai 200433, China
2. Department of Botany and Microbiology, University of Oklahoma, Norman, OK 73019, USA
3. Department of Environmental Science and Engineering, Fudan University, 220 Handan Road, Shanghai 200433, China