1. School of Agriculture, Food, & Wine and The Waite Research Institute, University of Adelaide, Waite Campus, PMB1, Glen Osmond, South Australia 5064, Australia
2. Present address: European Molecular Biology Laboratory, Grenoble Outstation, 38042 Grenoble, France
3. Australian Centre for Plant Functional Genomics, Waite Campus, PMB1, Glen Osmond, South Australia 5064, Australia