1. Key Laboratory of Conservation Biology for Endangered Wildlife of the Ministry of Education, and Laboratory of Systematic & Evolutionary Botany and Biodiversity, College of Life Sciences, Zhejiang University, Hangzhou 310058, China
2. Department of Organismic Biology, Salzburg University, A‐5020 Salzburg, Austria
3. Division of Forest and Biomaterials Science, Graduate School of Agriculture, Kyoto University, Kyoto 6068502, Japan
4. Department of Plant Ecology, Forestry and Forest Products Research Institute, Matsunosato, Tsukuba, Ibaraki 305‐8687, Japan
5. Field Centre for Sustainable Agriculture and Forestry, Faculty of Agriculture, Niigata University, 94‐2 Koda, Sado, Niigata 952‐2206, Japan