1. Department of Cell & Systems Biology and the Centre for the Analysis of Genome Evolution & Function, University of Toronto, Toronto, Ontario, Canada M5S 3B2
2. Department of Molecular & Cellular Biology, University of Guelph, Guelph, Ontario, Canada N1G 2W1
3. Program in Molecular Structure and Function, Hospital for Sick Children, College Street, Toronto, Ontario, Canada M5G 1L7