1. Center for Applied Conservation Research, Forest Sciences Centre; University of British Columbia; Vancouver BC V6T 1Z4 Canada
2. Department of Zoology and Faculty of Land and Food Systems; University of British Columbia; 6270 University Boulevard Vancouver BC V6T 1Z4 Canada
3. Fish Ecology and Conservation Physiology Laboratory, Ottawa-Carleton Institute for Biology; Carleton University; 1125 Colonel By Drive Ottawa Ontario K1S 5B6 Canada
4. Institute of Environmental Science; Carleton University; 1125 Colonel By Drive Ottawa Ontario K1S 5B6 Canada