1. Department of Biological Sciences; University of Alberta; Edmonton AB T6G 2E9 Canada
2. Department of Biology; Pontificia Universidad Javeriana; Carrera 7 No. 40-62 Bogotá D.C. Colombia
3. CNRS; UMR 5554 Institut des Sciences de l'Evolution (Université de Montpellier); Place Eugène Bataillon 34095 Montpellier France
4. Australian National Insect Collection; CSIRO National Research Collections Australia; Canberra ACT 2601 Australia