1. INRA; UMR1202 BIOGECO; F-33610 Cestas France
2. University of Bordeaux; BIOGECO; UMR1202; F-33400 Talence France
3. INRA; UMR1201 DYNAFOR; F-31326 Castanet-Tolosan France
4. Scion (New Zealand Forest Research Institute); P.O. Box 29237 Christchurch 8540 New Zealand