1. Institute of Biological and Environmental Sciences; University of Aberdeen; 23 St Machar Drive Aberdeen AB24 3UU UK
2. School of Ocean Sciences; University of Bangor; Menai Bridge Anglesey LL59 5AB UK
3. Biomathematics and Statistics Scotland; Craigiebuckler; Aberdeen AB15 8QH UK
4. Centre for Ecology & Hydrology; Bush Estate; Penicuik Midlothian EH26 0QB UK