1. CEFE, Univ Montpellier CNRS EPHE IRD Univ Paul Valéry Montpellier 3Montpellier France
2. UMR MIVEGEC Univ. MontpellierCNRSIRD Montpellier France
3. Department of Biology Utah State University Logan UT USA
4. Department of Biological Sciences University of Notre Dame Notre Dame IN USA