1. Division of Digestive Diseases, Department of Metabolism, Digestion and Reproduction St Mary's Hospital Campus, Imperial College London London UK
2. Departments of Gastroenterology and Hepatology St Mary's Hospital, Imperial College Healthcare NHS Trust London UK
3. Section of Bioinformatics, Division of Systems Medicine, Department of Metabolism, Digestion and Reproduction, Faculty of Medicine Imperial College London London UK
4. Division of Gastroenterology, Hepatology, and Endoscopy Brigham and Women's Hospital Boston Massachusetts USA
5. Harvard Medical School Boston Massachusetts USA