1. Department of Biology, Faculty of SciencePrince of Songkla University Hat Yai Songkhla Thailand
2. Seaweed and Seagrass Research Unit, Excellence Centre for Biodiversity of Peninsular Thailand, Faculty of SciencePrince of Songkla University Hat Yai Songkhla Thailand
3. Department of Biological ScienceSungkyunkwan University Suwon South Korea