1. Department of Genetics, Evolution and Environment; University College London; Darwin Building Gower Street London WC1E 6BT UK
2. School of Biological and Chemical Sciences; Queen Mary University of London; Mile End Road London E1 4NS UK
3. Center of Integrative Conservation; Xishuangbanna Tropical Botanical Garden; Chinese Academy of Sciences; Menglun Yunnan China
4. Department of Botany; Natural History Museum; Cromwell Road London SW7 5BD UK
5. School of Earth Sciences; University of Bristol; Life Sciences Building Tyndall Avenue Bristol BS8 1TQ UK