1. Centre for Neurogenetics and Statistical Genomics, Queensland Brain InstituteUniversity of Queensland St Lucia, Brisbane, Queensland 4072 Australia
2. School of Mathematics and PhysicsUniversity of Queensland St Lucia, Brisbane, Queensland 4072 Australia
3. Queensland Department of Agriculture and FisheriesEcosciences Precinct Joe Baker St, Dutton Park, Brisbane, Queensland 4102 Australia
4. School of Mathematical SciencesQueensland University of Technology Brisbane, Queensland 4000 Australia