1. Senckenberg Biodiversity and Climate Research Centre (BiK-F); Frankfurt am Main Germany
2. Institute for Ecology, Evolution and Diversity; Goethe University, Biologicum; Frankfurt am Main Germany
3. Department of Plant Systematics; University of Bayreuth; Bayreuth Germany
4. Department of Zoology and Wildlife Conservation; University of Dar es Salaam; Dar es Salaam Tanzania
5. Zoological Research Museum Alexander Koenig; Bonn Germany