1. Institut de Biologie Moléculaire des Plantes du CNRS; Université de Strasbourg; 12 rue du Général Zimmer Strasbourg 67084 France
2. State Key Laboratory of Genetic Engineering; Collaborative Innovation Center for Genetics and Development; International Associated Laboratory of CNRS-Fudan-HUNAU on Plant Epigenome Research; Department of Biochemistry; Institute of Plant Biology; School of Life Sciences; Fudan University; Shanghai 20043 China