1. School of Ocean Sciences College of Natural Sciences Bangor University Menai Bridge LL59 5AB UK
2. Department of Fish, Wildlife and Conservation Biology Colorado State University Fort Collins CO 80523 USA
3. Institute of Biological and Environmental Sciences University of Aberdeen Lighthouse Field Station George Street Cromarty IV11 8YJ UK
4. Carrigskeewaun Carrowniskey Westport, Co. Mayo Ireland
5. Coolmore Rossnowlagh, Co. Donegal Ireland