1. Centre de Biologie pour la Gestion des Populations (CBGP); UMR INRA-IRD-CIRAD-Montpellier SupAgro; Campus International de Baillarguet; 34988 Montferrier-sur-Lez Cedex France
2. UMR 5174 EDB (Laboratoire Evolution & Diversité Biologique); Université de Toulouse; ENFA; 118 route de Narbonne F-31062 Toulouse France
3. UMR 5174 EDB; CNRS; Université Paul Sabatier; F-31062 Toulouse France
4. State Key Laboratory for the Biology of the Plant Diseases and Insect Pests; Institute of Plant Protection; Chinese Academy of Agricultural Sciences; No.2 West Yuanmingyuan Road Beijing 100193 China
5. UMR 5175; CNRS; Centre d'Ecologie Fonctionnelle et Evolutive (CEFE); 1919 route de Mende Montpellier Cedex 05 34293 Montpellier France
6. School of biological technology; Xi'an University of Arts and Science; No.168 South Taibai Road Xi'an Shaanxi Province 710065 China