1. Graduate School of Agriculture Kyoto University, Kitashirakawa Oiwake‐cho, Sakyo‐ku Kyoto 606‐8502 Japan
2. Graduate School of Life and Environmental Sciences Osaka Prefecture University, 1‐1, Gakuen‐cho, Naka‐ku, Sakai Osaka 599‐8531 Japan
3. Graduate School of Agricultural Science Tohoku University, 232‐3, Yomogida, Naruko‐onsen, Osaki Miyagi 989‐6711 Japan