1. Division of Applied Life Science (BK21 Program), IALS, PMBBRC, Gyeongsang National University; Jinju; 660-701; Korea
2. Korea Research Institute of Bioscience and Biotechnology; 52 Eoeun-dong, Yuseong-gu; Daejon; 305-806; Korea
3. Department of Environmental Horticulture, University of Seoul; Seoul; 130-743; Korea
4. Institute of Botany, The Chinese Academy of Sciences; Beijing; 100093; China