1. Department of Plant Sciences; University of Cambridge; Downing Street; Cambridge; CB2 3EA; U.K.
2. Landcare Research; PO Box 40; Lincoln; 7640; New Zealand
3. Faculty of Environmental Earth Science; Hokkaido University; Sapporo; 060-0810; Japan
4. Department of Integrative Ecology; Estación Biológica de Doñana (EBD-CSIC); C/Américo Vespucio s/n; Sevilla; 41092; Spain