1. Division of Agricultural and Environmental Sciences; School of Biosciences; University of Nottingham; Sutton Bonington Campus Loughborough Leicestershire LE12 5RD UK
2. Department of Sustainable Soils and Grassland Systems; Rothamsted Research; Harpenden Hertfordshire AL5 2JQ UK
3. Centre for Environmental Geochemistry; Inorganic Geochemistry; British Geological Survey; Nicker Hill Keyworth Nottingham NG12 5GG UK