1. Department of Biological and Environmental Sciences; University of Gothenburg; Box 461 SE-405 30 Göteborg Sweden
2. Department of Biological Sciences; University of Alberta; Edmonton T6G 2E9 AB Canada
3. CNRS, UMR 5554 Institut des Sciences de l'Evolution, Université de Montpellier; Place Eugène Bataillon 34095 Montpellier France
4. Department of Ecology and Evolutionary Biology; Brown University; Providence RI 02912 USA
5. Gothenburg Botanical Garden; Carl Skottsbergs gata 22A 413 19 Gothenburg Sweden