1. Section of Animal Biology; Department of Biology; University of Patras; Patras GR-26504 Greece
2. Centre for Ecology & Hydrology; Penicuik EH26 0QB UK
3. Department of Mathematics and Geosciences; University of Trieste; I-34127 Trieste Italy
4. Institute of Biological and Environmental Sciences; University of Aberdeen; Aberdeen AB24 2TZ UK