1. Department of Animal Development and Physiology; Graduate School of Biostudies, Kyoto University; Kyoto; 606-8501; Japan
2. Department of Developmental Biology; National Institute for Basic Biology; Okazaki; 444-8585; Japan
3. Science Center, Tensei-suisan Co., Ltd; Karatsu; 847-0193; Japan
4. Drug Discovery Research Laboratories; Kyowa Hakko Kirin Co., Ltd; Shizuoka; 411-8731; Japan
5. Department of Cell Biology; Research Institute for Microbial Diseases, Osaka University; Suita; 565-0871; Japan
6. Department of Zoology; Graduate School of Science, Kyoto University; Kyoto; 606-8502; Japan