1. Department of Ecology and Evolutionary Biology University of Michigan Ann Arbor Michigan USA
2. Department of Biology Oxford USA
3. School of Natural and Environmental Sciences Newcastle University Newcastle‐upon‐Tyne UK
4. Laboratoire de Biométrie et Biologie Evolutive Université de Lyon, Université Lyon 1, CNRS Villeurbanne France
5. School of Biological Sciences University of Aberdeen Aberdeen UK