1. Department of Biological Sciences; Marine Sciences Program; Florida International University; North Miami FL 33181 USA
2. Southeast Environmental Research Center; Florida International University; Miami FL 33199 USA
3. School of Environmental and Forest Sciences; University of Washington; Seattle WA 98195 USA
4. Evolutionary and Behavioural Ecology Research Group; Department of Biological Sciences; Simon Fraser University; Burnaby BC V5A 1S6 Canada