1. Leibniz-Institute of Freshwater Ecology and Inland Fisheries; Mäggelseedamm 310 12587 Berlin Germany
2. Institute of Biology; Freie Universität Berlin; Schwendenerstr. 1 14195 Berlin Germany
3. Theoretical Evolutionary Ecology; Department of Animal Ecology and Tropical Biology; Biocenter; Emil-Fischer-Str. 32 97074 Würzburg Germany
4. Biology Department; Willamette University; 900 State Street Salem OR 97301 USA