1. Département de biologie; Université de Sherbrooke; Sherbrooke Québec J1K 2R1 Canada
2. Department of Mathematics and Computer Sciences; IMADA; University of Southern Denmark; Odense Denmark
3. Max-Planck Odense Center on the Biodemography of Aging; MaxO; University of Southern Denmark; Odense Denmark
4. Departamento de Conservación de la Biodiversidad; El Colegio de la Frontera Sur; Chetumal Mexico