1. UWA School of Agriculture and Environment, Faculty of ScienceThe University of Western Australia Crawley WA 6009 Australia
2. Graduate School of Bioagricultural SciencesNagoya University Nagoya 464‐8601 Japan
3. Freshwater Biological Laboratory, Department of BiologyUniversity of Copenhagen 2100 Copenhagen Denmark
4. Aarhus Institute of Advanced StudiesAarhus University 8000 Aarhus Denmark