1. Department of Biology, Integrated Bioscience Program; University of Akron; Akron Ohio 44325
2. Department of Ecology, Evolution, and Environmental Biology; Columbia University; New York New York 10027
3. Center for Integrative Animal Behavior; Columbia University; New York New York 10027
4. Department of Biology, Terrestrial Ecology Unit; University of Ghent; Ledeganckstraat 35 9000 Ghent Belgium