1. Department of Gastroenterology and Metabolism Graduate School of Biomedical and Health Sciences Hiroshima Japan
2. Research Center for Hepatology and Gastroenterology Hiroshima University Hiroshima Japan
3. Department of Epidemiology Infectious Disease Control and Prevention Graduate school of Biomedical and Health Sciences Hiroshima University Hiroshima Japan
4. Collaborative Research Laboratory of Medical Innovation Hiroshima University Hiroshima Japan
5. RIKEN Center for Integrative Medical Sciences Yokohama Japan