1. CAS Key Laboratory of Tropical Forest Ecology, Xishuangbanna Tropical Botanical Garden Chinese Academy of Sciences Menglun Yunnan 666303 China
2. Center of Conservation Biology, Core Botanical Gardens Chinese Academy of Sciences Menglun Yunnan 666303 China
3. Experimental Center of Tropical Forestry Chinese Academy of Forestry Pingxiang Guangxi Zhuang Autonomous Region 532600 China
4. Department of Ecology and Environmental Science, UPSC Umeå University Umeå 90187 Sweden
5. Center for Integrative Conservation, Xishuangbanna Tropical Botanical Garden Chinese Academy of Sciences Menglun Yunnan 666303 China
6. State Key Laboratory of Tree Genetics and Breeding Chinese Academy of Forestry Beijing 100091 China