1. School of Agriculture and Food Sciences; The University of Queensland; St. Lucia Qld 4072 Australia
2. Biosecurity Queensland, Department of Agriculture and Fisheries, Ecosciences Precinct; Brisbane Qld 4001 Australia
3. Department of Primary Industry and Resources; Northern Territory Government; Darwin NT 0801 Australia
4. School of Plant Biology; The University of Western Australia; Crawley WA 6009 Australia