1. The Hawkesbury Institute for the Environment Western Sydney University Richmond New South Wales Australia
2. Research School of Biology The Australian National University Canberra Australian Capital Territory Australia
3. Centre for Sustainable Ecosystem Solutions, School of Earth, Atmospheric and Life Sciences University of Wollongong Wollongong New South Wales Australia
4. Fuller Evolutionary Biology Program, Cornell Lab of Ornithology Cornell University Ithaca New York USA