1. Frankfurt Zoological Society, Selous Wild Dog Project, c/o Selous Conservation Programme, Box 1519, Dar es Salaam, Tanzania
2. *Field Research Center for Ecology and Ethology, Rockefeller University, RR2 Box 38B, Tyrrel Road, Millbrook, New York 12545, USA
3. Sokoine University of Agriculture, Faculty of Veterinary Medicine, Box 3021, Morogoro, Tanzania
4. Durrell Institute of Conservation and Ecology, The University, Canterbury, Kent CT2 7NX, UK