1. Evolutionary Genomics Group, Department of Botany and Zoology, Stellenbosch University, Matieland, South Africa
2. Department of Botany and Zoology, Centre for Invasion Biology, Stellenbosch University, Matieland, South Africa
3. Centre for Veterinary Science, University of Cambridge, Cambridge, UK
4. The Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, Cambridge, UK