1. Department of Earth Sciences, University of Bristol, Bristol, UK
2. School of Biology and Environmental Sciences, University College Dublin, Belfield, Dublin, Ireland
3. Department of Anatomy, College of Osteopathic Medicine of the Pacific and College of Podiatric Medicine, Western University of Health Sciences, Pomona, CA, USA
4. Palaeobiology Research Group, School of Earth and Environmental Sciences, University of Portsmouth, Portsmouth, UK