1. Montana Cooperative Wildlife Research Unit Wildlife Biology Program University of Montana Missoula MT USA
2. Montana Cooperative Wildlife Research Unit Division of Biological Sciences University of Montana Missoula MT USA
3. FitzPatrick Institute of African Ornithology Department of Science and Technology/National Research Foundation Centre of Excellence University of Cape Town Private Bag X3, Rondebosch 7701 South Africa
4. Department of Biology University of New Mexico Albuquerque NM USA
5. U.S. Geological Survey Montana Cooperative Wildlife Research Unit University of Montana Missoula MT USA