1. Ministry of Education Key Laboratory for Ecosecurity of Southwest China, Yunnan Key Laboratory of Plant Reproductive Adaptation and Evolutionary Ecology and Centre for Invasion Biology Institute of Biodiversity, School of Ecology and Environmental Science, Yunnan University Kunming Yunnan China
2. Key Laboratory of National Forestry & Grassland Administration on Forest Resources Conservation and Ecological Safety in the Upper Reaches of the Yangtze River College of Forestry, Sichuan Agricultural University Chengdu China
3. College of Agriculture Sichuan Agricultural University Chengdu China
4. Above and Belowground Interactions, Institute of Biology Leiden University Leiden The Netherlands