1. Plant Conservation and Population Biology; University of Leuven; Kasteelpark Arenberg 31-2435 BE-3001 Leuven Belgium
2. Department of Horticulture and Plant Science; Jimma University; PO Box 307 Jimma Ethiopia
3. Division Forest, Nature and Landscape; University of Leuven; Celestijnenlaan 200E-2411 BE-3001 Leuven Belgium
4. Plant Sciences Unit - Growth and Development; Institute for Agricultural and Fisheries Research; Caritasstraat 21 9090 Melle Belgium