1. Institute of Applied Ecology Chinese Academy of Sciences Shenyang China
2. University of Chinese Academy of Sciences Beijing China
3. Department of Biology Plant Conservation and Population Biology KU Leuven Leuven Belgium
4. Shenyang ArboretumChinese Academy of Sciences Shenyang China
5. Key Laboratory of Forest Ecology and Management Chinese Academy of Sciences Shenyang China
6. Meise Botanic Garden Meise Belgium