1. Department of Biology University of Puerto Rico‐Río Piedras PO Box 23360 San Juan PR 00936‐3360 USA
2. International Institute of Tropical Forestry United States Forest Service United States Department of Agriculture Jardín Botánico Sur, 1201 Calle Ceiba San Juan PR 00926 USA
3. Center for Applied Tropical Ecology and Conservation Department of Biology University of Puerto Rico PO Box 23360 San Juan PR 00936‐3360 USA