1. Atmosphere and Ocean Research Institute The University of Tokyo Kashiwa Chiba Japan
2. Department of Biological Sciences, Graduate School of Science The University of Tokyo Bunkyo‐ku Tokyo Japan
3. Graduate School of Fisheries Sciences Hokkaido University Hakodate Hokkaido Japan
4. Division of Applied Bioscience, Research Faculty of Agriculture Hokkaido University Sapporo Hokkaido Japan