1. National Centre for Biosystematics, Natural History Museum, University of Oslo, P.O.Box 1172 Blindern, NO‐0318 Oslo, Norway
2. Laboratoire d’Ecologie Alpine, UMR UJF‐CNRS 5553, Université Joseph Fourier, 38041 Grenoble, France
3. Centre for Ecological and Evolutionary Synthesis, Department of Biology, University of Oslo, P.O.Box 1066 Blindern, NO‐0316 Oslo, Norway
4. School of Geography, University of Southampton, Room 2019 Shackleton Building, Highfield, Southampton, SO17 1BJ, UK
5. Centre for Ancient Genetics, University of Copenhagen, Universitetsparken 15, DK‐2100, Denmark
6. Department of Zoology, University of Oxford, South Parks Road, Oxford OX1 3PS, UK